I've been researching all this recently for a book. My feeling is that lunar+solar calendars are recent in comparison with the length of Ice Age human existence, e.g. Warren Field, which is about 8000BC. Hunter-gatherers (really, we should be calling them gatherer-hunters, as the former food source for the vast majority of prehistoric peoples is much more important) would certainly have recognised the Moon as a celestial timekeeper. They wouldn't particularly have needed to know about equinoxes, but they did need a general comprehension of natural history, into which midwinter, midsummer etc would have fed.
If you look for instance at the Lascaux cave paintings, the three main species depicted, aurochs bull, deer and horse, are all depicted how they appear in their mating season. So there would have been generalised connections in their minds, passed on orally down the generations, between natural events and themselves.