Well, this is a thorny one!
In Middle-earth, it appears to be Sauron, (the Olog-Hai):
Is this what you had in mind? or do I need to dig more?
In Middle-earth, it appears to be Sauron, (the Olog-Hai):
LotR - Appendix F - Of Other Races."That Sauron bred them none doubted, though from what stock was not known."
Is this what you had in mind? or do I need to dig more?